Western Barking Frog Craugastor augusti cactorum (Taylor, 1938)
An adult specimen from Santa Cruz County, Arizona Image © Suzanne L. Collins, 2004
Taxonomic Comments:
Caren S. Goldberg, Brian K. Sullivan, John H. Malone and Cecil R. Schwalbe (2004. Divergence among Barking Frogs (Eleutherodactylus augusti) in the Southwestern United States. Herpetologica 60(3): 312-320), using call variation and mtDNA, demonstrated that the U.S. subspecies, E. a. cactorum, may be a distinct species. Additional data is needed from Mexican specimens to clarify the status of this taxon.